[1] [1] PULLIN J, BRADLEY J M, MOORE G R, et al. Electron transfer from haem to the di-iron ferroxidase centre in bacterioferritin［J］. Angewandte Chemie-International Edition, 2021, 60(15):8376-8379.

[2] [2] HE D, PIERGENTILI C, ROSS J, et al. Conservation of the structural and functional architecture of encapsulated ferritins in bacteria and archaea［J］. Biochemical Journal, 2019, 476(6): 975-989.

[3] [3] WANG W M, LIU X J, WANG Y J, et al. Distinct structural characteristics define a new subfamily of Mycoplasma ferritin［J］.Chinese Chemical Letters, 2022, 33(11): 4952-4955.

[4] [4] WANG W M, ZHANG Y, ZHAO G H, et al. Ferritin with atypicalferroxidase centers takes B-channels as the pathway for Fe2+ uptake from Mycoplasma［J］. Inorganic Chemistry, 2021, 60(10):7207-7216.

[5] [5] HOUSER K V, CHEN G L, CARTER C, et al. Safety and immunogenicity of a ferritin nanoparticle H2 influenza vaccine inhealthy adults: a phase 1 trial［J］. Nature Medicine, 2022, 28(2):383-391.

[6] [6] POWELL A E, ZHANG K M, SANYAL M, et al. A single immunization with spike-functionalized ferritin vaccines elicits neutralizing antibody responses against SARS-CoV-2 in mice［J］. ACSCentral Science, 2021, 7(1): 183-199.

[7] [7] KANEKIYO M, BU W, JOYCE M G, et al. Rational design of anEpstein-Barr virus vaccine targeting the receptor-binding site［J］.Cell, 2015, 162(5): 1090-1100.

[8] [8] WANG Wen, GU Zhifeng, ZHU Ningning, et al. Histopathology and ultrapathology of rickettsia-like organisms in Eriocheir sinensis with tremor disease［J］. Journal of Fishery Sciences ofChina, 2001, 8(4): 32-35, 105-106.

[9] [9] WANG Wen, ZHU Ningning, LI Zhengrong, et al. Optical andtransmission electron microscope study on nerve tissue infestedby Rickettsia-like organisms in crab, Eriocheir sinensis, suffering from tremor disease［J］. Zoological Research, 2001, 22(6):467-471, 523.

[10] [10] WANG Wen. Discovery and studies of a novel pathogen, Spiroplasma eriocheiris, in economically important species of crustaceans［J］. Journal of Nanjing Normal University (Natural ScienceEdition), 2016, 39(1): 1-13.

[11] [11] LIANG Tingming. Screening and research of important functionalgenes of Spiroplasma in Eriocheir sinensis and multiplex PCRdetection technology of Spiroplasma and WSSV［D］. Nanjing:Nanjing Normal University, 2012.

[12] [12] CHEN R, ZHAO L, GAN R, et al. Evidence for the rapid anddivergent evolution of Mycoplasmas: structural and phylogeneticanalysis of enolases［J］. Frontiers in Molecular Biosciences,2021, 8: 811106.

[13] [13] OTWINOWSKI Z, MINOR W. Processing of X-ray diffractiondata collected in oscillation mode［J］. Methods in Enzymology,1997, 8(276): 307-326.

[14] [14] CHEN R, YU Y, FENG Z, et al. Featured species-specific loopsare found in the crystal structure of Mhp Eno, a cell surface adhesin from Mycoplasma hyopneumoniae［J］. Frontiers in Cellularand Infection Microbiology, 2019, 9(209): 1-14.

[15] [15] BLAESI E J, PALOWITCH G M, HU K, et al. Metal-free classIe ribonucleotide reductase from pathogens initiates catalysis witha tyrosine-derived dihydroxyphenylalanine radical［J］. Proceedings of the National Academy of Sciences of the United States ofAmerica, 2018, 115(40): 10022-10027.

[16] [16] SRINIVAS V, LEBRETTE H, LUNDIN D, et al. Metal-free ribonucleotide reduction powered by a DOPA radical in Mycoplasma pathogens［J］. Nature, 2018, 563(7731): 416-420.

[17] [17] MASUDA T, MORIMOTO S, MIKAMI B, et al. The extensionpeptide of plant ferritin from sea lettuce contributes to shell stability and surface hydrophobicity［J］. Protein Science, 2012, 21(6):786-796.

[18] [18] MARCHETTI A, PARKER M S, MOCCIA L P, et al. Ferritin isused for iron storage in bloom-forming marine pennate diatoms［J］. Nature, 2009, 457(7228): 467-470.

[19] [19] SUBHADARSHANEE B, MOHANTY A, JAGDEV M K, et al. Surface charge dependent separation of modified and hybridferritin in native PAGE: impact of lysine 104［J］. Biochimicaet Biophysica Acta-Proteins and Proteomics, 2017, 1865(10):1267-1273.

[20] [20] DE MEULENAERE E, BAILEY J B, TEZCAN F A, et al. Firstbiochemical and crystallographic characterization of a fastperforming ferritin from a marine invertebrate［J］. BiochemicalJournal, 2017, 474(24): 4193-4206.

[21] [21] HAMBURGER A E, WEST A P, HAMBURGER Z A, et al. Crystal structure of a secreted insect ferritin reveals a symmetricalarrangement of heavy and light chains［J］. Journal of MolecularBiology 2005, 349(3): 558-569.

[22] [22] JOHNSON E, CASCIO D, SAWAYA M R, et al. Crystal structuresof a tetrahedral open pore ferritin from the hyperthermophilicarchaeon Archaeoglobus fulgidus［J］. Structure, 2005, 13(4):637-648.