[1] [1] Chen, C., He, G., Li, J., Perez-Hormaeche, J., Becker, T., Luo, M., Wallrad, L., Gao, J., Li, J., Pardo, J.M., et al.(2023). A salt stress-activated GSO1-SOS2-SOS1 module protects theArabidopsisroot stem cell niche by enhancing sodium ion extrusion. EMBO J.42:e113004.

[2] [2] Gao, Y.-Q., Huang, J.-Q., Reyt, G., Song, T., Love, A., Tiemessen, D., Xue, P.-Y., Wu, W.-K., George, M.W., Chen, X.-Y., et al.(2023). A dirigent protein complex directs lignin polymerization and assembly of the root diffusion barrier. Science382:464-471.

[3] [3] Gao, Y.-Q., Su, Y., and Chao, D.-Y.(2024). Exploring the function of plant root diffusion barriers in sealing and shielding for environmental adaptation. Nat. Plants10:1865-1874.

[4] [4] Pfister, A., Barberon, M., Alassimone, J., Kalmbach, L., Lee, Y., Vermeer, J.E.M., Yamazaki, M., Li, G., Maurel, C., Takano, J., et al.(2014). A receptor-like kinase mutant with absent endodermal diffusion barrier displays selective nutrient homeostasis defects. eLife3:e03115.

[5] [5] Salas-Gonzlez, I., Reyt, G., Flis, P., Custdio, V., Gopaulchan, D., Bakhoum, N., Dew, T.P., Suresh, K., Franke, R.B., Dangl, J.L., et al.(2021). Coordination between microbiota and root endodermis supports plant mineral nutrient homeostasis. Science371:eabd0695.

[6] [6] Shen, D., Wippel, K., Remmel, S., Zhang, Y., Kuertoes, N., Neumann, U., Kopriva, S., and Andersen, T.G.(2024). TheArabidopsisSGN3/GSO1 receptor kinase integrates soil nitrogen status into shoot development. EMBO J.43:2486-2505.

[7] [7] Shen, D., Micic, N., Venado, R.E., Bjarnholt, N., Crocoll, C., Persson, D.P., Samwald, S., Kopriva, S., Westhoff, P., Metzger, S., et al.(2025). Apoplastic barriers are essential for nodule formation and nitrogen fixation inLotus japonicus. Science387:1281-1286.

[8] [8] Song, T., Tian, Y.-Q., Liu, C.-B., Gao, Y.-Q., Wang, Y.-L., Zhang, J., Su, Y., Xu, L.-N., Han, M.-L., Salt, D.E., and Chao, D.Y.(2023). A new family of proteins is required for tethering of Casparian strip membrane domain and nutrient homoeostasis in rice. Nat. Plants9:1749-1759.

[9] [9] Verbon, E.H., Liberman, L.M., Zhou, J., Yin, J., Pieterse, C.M.J., Benfey, P.N., Stringlis, I.A., and de Jonge, R.(2023). Cell-type-specific transcriptomics reveals that root hairs and endodermal barriers play important roles in beneficial plant-rhizobacterium interactions. Mol. Plant16:1160-1177.

[10] [10] Wang, F.-L., Tan, Y.-L., Wallrad, L., Du, X.-Q., Eickelkamp, A., Wang, Z.-F., He, G.-F., Rehms, F., Li, Z., Han, J.-P., et al.(2021). A potassium-sensing niche inArabidopsisroots orchestrates signaling and adaptation responses to maintain nutrient homeostasis. Dev. Cell56:781-794.e6.